ARTIFICIAL KEY TO THE FOREGOING HIGHER GROUPS,
By means of which any North American bird may be readily referred to that group to which it is held to belong.
I. Toes 3; 2 in front, 1 behind (Pici) Picariæ.
II. Toes 3; all in front.
Toes cleft or semipalmate Limicolæ.
Toes palmate.
Nostrils tubular Longipennes.
Nostrils not tubular Pygopodes.
III. Toes 4; 2 in front, 2 behind.
Bill cered and hooked Psittaci.
Bill neither cered nor hooked. (Cuculi or Pici) Picariæ.
IV. Toes 4; 3 in front, 1 behind.
1. Toes syndactyle (Cuculi) Picariæ.
2. Toes totipalmate (all four full-webbed) Steganopodes.
3. Toes palmate.
Bill curved up Limicolæ.
Bill not curved up;
lamellate Lamellirostres.
not lamellate;
hallux lobate Pygopodes.
hallux not lobate Longipennes.
4. Toes lobate.
Tail rudimentary Pygopodes.
Tail perfect.
A horny frontal shield Alectorides.
No horny frontal shield Limicolæ.
5. Toes semipalmate;
joined by evident movable basal web (A).
6. Toes cleft to the base,
or there immovably coherent (B).
A. Hind toe elevated above the level of the rest.
Tibiæ naked below.
Nostrils perforate Alectorides.
Nostrils imperforate.
Tarsi reticulate.
Head bald Herodiones.
Head feathered Limicolæ.
Tarsi scutellate in front Limicolæ.
Tibiæ feathered below.
Nostrils perforate Raptores.
Nostrils imperforate.
Gape reaching below eye. (Cypseli) Picariæ.
Gape not reaching below eye Gallinæ.
AA. Hind toe inserted on the level of the rest.
Tibiæ naked below Herodiones.
Tibiæ feathered below.
Bill cered and hooked Raptores.
Bill not cered.
Nasal membrane soft and tumid Columbæ.
Nasal scale hard and flat Gallinæ.
B. Hind toe elevated above the level of the rest.
Gape reaching below eye (Cypseli) Picariæ.
Gape not below eye.
First primary emarginate
or about equal to 2d Limicolæ.
First primary not emarginate
and much shorter than 2d Alectorides.
BB. Hind toe inserted on the level of the rest.
Nostrils opening beneath soft swollen membrane Columbæ.
Nostrils otherwise.
Bill cered and hooked Raptores.
Bill otherwise.
Secondaries only six (Cypseli) Picariæ.
Secondaries more than six (a) Passeres.
(a) Primaries 10; the 1st
more than 2⁄3 as long as the longest. (Clamatores) Passeres.
Primaries 10; the 1st
not 2⁄3 as long as the longest. (Oscines) Passeres.
Primaries 9. (Oscines) Passeres.
Recurring now to consideration of the North American Families of the foregoing higher groups, I take up the latter in the natural order in which they have been presented, giving under head of each such group an analysis of the North American families by which it is represented, reiterating the caution that the characters are drawn up only with reference to the North American genera, and are, consequently, not necessarily or always applicable upon wider considerations. These analyses are made as nearly natural as the state of the case permits, but I seize upon any obvious external characters which may be afforded, without regard to their morphological significance or taxonomic value.
Analysis of the Families of PASSERES
A. Oscines. Musical apparatus highly developed. Back of tarsus undivided, or formed of a few scutella distinct from those lapping over the front. First primary wanting, spurious, or at most not over two thirds the length of the longest.
a. Each side of tarsus covered with a plate undivided in most or all of its length, and meeting its fellow in a sharp ridge behind.
b. Primaries only nine.
c. Bill triangular, depressed, about as wide at base as long; the gape twice as long as the culmen, reaching to about opposite the eyes; tomia straight or gently curved. No obvious rictal bristles. Tarsi not longer than the lateral toe and claw. Wings long and pointed, the first primary equal to or longer than the second. Central tail-feathers not half as long as the wing … Hirundinidæ.
cc. Bill variously conico-elongate or slender, or, if depressed, with long rictal bristles; gape not nearly twice as long as culmen; tomia straight or gently curved. Nostrils not obviously nearer culmen than tomia. Tarsus longer than lateral toe and claw.
Bill very slender, acute; culmen rather concave at base. Longest secondary acuminate, nearly or quite equal to the primaries in the closed wing. Hind claw little curved, about twice as long as the middle claw. Hind toe and claw longer than middle toe and claw … Motacillidæ.
Bill variously conico-elongate and acute; culmen not concave at base. Longest secondary not acuminate, falling far short of primaries in the closed wing. Hind claw well curved, not nearly twice as long as middle claw; hind toe and claw not longer than middle toe and claw. Gape ample; tongue slightly bifid or brushy, if at all … Sylvicolidæ.
Bill lengthened, very acute, even decurved. Wings and feet as in the last. Gape constricted; tongue generally deeply bifid or brushy … Cærebidæ.
ccc. Bill more or less truly conic, usually short, thick; commissure usually more or less evidently abruptly angulated near the base, or with lobe or tooth further forward. Nostrils obviously nearer culmen than tomia. Tarsus longer than lateral toe and claw.13
Bill stout, tumid, convex in nearly all its outlines; tomia not angulated, but with one or more lobes or nicks in advance of the base. Nostrils placed very high. Other characters much as in Sylvicolidæ. Colors chiefly red and yellow. One genus of … Tanagridæ.
Bill truly conic, much shorter than head, usually with the angulation evident; no lobe along middle of tomia, but usually a notch at end. Nostrils placed very high. Rictal bristles usually obvious … Fringillidæ.
Bill conic, but lengthened, little if any shorter than head; the angulation of the tomia evident; no notch at end. Nostrils high. No rictal bristles … Icteridæ.
bb. Primaries ten.
Otherwise with characters much as in Icteridæ … Sturnidæ.
d. Nostrils concealed with antrorse bristly feathers (except in Psilorhinus and Gymnokitta).14
Base of bill sheathed with antrorse bristly feathers, having lateral branches to their very ends; its tip mostly notched. Basal joint of middle toe united only half-way to the lateral. Sides of tarsus occupied by a lateral groove, mostly filled in with small plates. First primary more than half as long as second. Large,—over seven inches … Corvidæ.
Base of bill with two tufts of bristly feathers, ending in simple filaments without lateral branches, its tip mostly unnotched. Basal joint of middle toe united nearly all its length with the lateral. Sides of tarsus ungrooved. First primary less than half as long as second. Small,—under seven inches … Paridæ.15
dd. Nostrils exposed.
e. Tail scansorial, with rigid acute feathers. Whole bill slender, compressed, acute, decurved, unnotched, unbristled. Outer toe much longer than inner … Certhiidæ.
ee. Tail not scansorial, graduated. First primary not less (generally more) than half as long as the second, and inner toe united to the middle by at least one half (usually more) of the length of its basal joint.
Tarsus with few obscure scutella. Rictal bristles present. Bill stout, but not toothed nor hooked. Wing excessively rounded (fifth, sixth, and seventh primaries longest), much shorter than the long graduated tail. Size small. Plumage brown, unbanded … Chamæadæ.
Tarsus distinctly scutellate. Nostrils wholly exposed, scaled. No rictal bristles, but loral feathers with bristly points. Bill slender, not notched nor hooked. Wings and tail moderately rounded; neither very much shorter than the other. Size small. Color brown, etc., the wings and tail barred or undulated … Troglodytidæ.
Tarsus distinctly scutellate. Nostrils overhung (not concealed) with bristly feathers. Rictal bristles present, strong. Bill powerful, compressed, strongly notched, toothed, and hooked. Wings and tail moderate. Large. Colors black, white, and gray … Laniidæ.
eee. Tail not scansorial. First primary less than half as long as the second,16 or about half as long, in which case the inner toe is cleft nearly to its base (f and ff).
f. Basal joint of middle toe united some distance with the inner, and for half or more of its length with the outer toe.
Basal joint of middle toe shorter than that of inner toe, and wholly adherent to both inner and outer toes. Tarsus longer than middle toe and claw. Gonys more than half the length of the lower jaw. Bill stout, high, compressed; notched and abruptly hooked at tip … Vireonidæ.
Basal joint of middle toe not shorter than that of inner toe; united to the outer for about two thirds, to the inner for about one half, its length. Tarsus not longer than the middle toe and claw. Gonys less than half the length of the under jaw. Bill triangular, much depressed at base, moderately notched, and hooked at tip17 … Ampelidæ.
Basal joint of middle toe shorter than that of the inner toe, united to the outer for about two thirds, to the inner for about one half, its length. Tarsus longer than middle toe and claw. Gonys more than half the length of the under jaw. Bill very weak and slender, little decurved or notched at tip. Very small,—under six inches long. (Tarsi booted in Regulus, distinctly scutellate in Polioptila.) … Sylviidæ.
ff. Basal joint of middle toe quite free from the inner, and not united with the outer more than half-way.
Nostrils linear, low. No bristles or bristly points whatever about the mouth. Wings short, rounded, concavo-convex. Tail very short, nearly concealed by its coverts. Tarsi booted … Cinclidæ.
Nostrils oval. Bristles or bristly points about the mouth. Wings very long and pointed, reaching, when folded, beyond the middle of the short, square, or emarginate tail, and one and a half times or more the length of the latter; tip formed by second, third, and fourth quills; outer secondary reaching only about two thirds way to end of longest primary; spurious quill very short. Tarsi booted … Saxicolidæ.
Nostrils oval. Bristles or bristly points about the mouth. Wings moderate, not reaching, when folded, beyond the middle of the tail, and not over one and a third times as long as the latter; tip formed by third to sixth quill; outer secondary reaching in closed wing three fourths or more the length of the longest primary. Spurious quill longer, sometimes one half the second. Tarsi scutellate in Miminæ, booted in Turdinæ … Turdidæ.
aa. Outside of tarsus covered with two series of scutella,—one lapping entirely around in front, the other entirely around behind, and meeting at a groove on the inside; hind edge blunt. First primary spurious or apparently wanting. Hind claw much lengthened, scarcely curved. Nostrils with antrorse bristly feathers. Bill conico-elongate … Alaudidæ.
B. Clamatores. Outside of tarsus covered with a series of plates variously arranged, lapping entirely around in front and behind, to meet at a groove on the inner side.
First primary lengthened, often longest, at least over two thirds as long as the longest. Bill broad at the base, much depressed, tapering to a fine point, which is abruptly decurved; culmen rounded or flattened; gonys flattened; commissure straight, or nearly so, to the tip. Nostrils small, circular, basal; overhung, but not concealed by bristles. Mouth capacious, with broad and deeply fissured rictus, beset with numerous long strong bristles. Feet small, weak. Tail of twelve feathers … Tyrannidæ.
Analysis of the Families of PICARIÆ
Secondaries only six.
Bill tenuirostral, longer than head, nearly cylindrical. Gape constricted. Tongue filiform, extensile, bi-tubular. Wings long in terminal portion, abbreviated proximally, acute. Plumage compact, of metallic sheen. Size smallest of all birds. (Humming-Birds.) … Trochilidæ.
Secondaries more than six.
Feet syndactyle by connation of outer and middle toes.
Outer toe much longer than the inner, united for half its length with the middle, forming a broad sole. Tibiæ naked below. Bill longer than head, straight, acute, with hard cutting edges and ample rictus. Tongue rudimentary, fixed. Wings pointed, much longer than the short square tail. Tail-feathers twelve. Plumage compact, oily. (Kingfishers.) … Alcedinidæ.
Feet zygodactyle18 by reversion of outer or fourth toe.
Not scansorial; tail of eight or ten long soft feathers. Bill with decurved tip, not fitted for hammering; rictus ample. Tongue not extensile nor vermiform nor barbed. Salivary glands and hyoidean apparatus not peculiar. No nasal tufts of feathers. Arboreal and terrestrial. (Cuckoos.) … Cuculidæ.
Highly scansorial; tail of twelve rigid acuminate feathers, whereof the outer pair are short and spurious, concealed between bases of next two pairs. Bill stout, straight, with the tip truncate or acute, not decurved,—an efficient chisel for hammering and boring wood. Tongue vermiform, extensile,19 and barbed. Salivary glands large; hyoidean apparatus peculiar. Nasal tufts usually present. Arboreal. (Woodpeckers.) … Picidæ.
Feet neither syndactyle nor zygodactyle.
Feet semipalmate, of normal ratio of phalanges. Anterior toes connected at base by movable webbing. Hind toe very small, elevated, semilateral. Middle toe produced, its large claw pectinate. Bill fissirostral, with very small, triangular, depressed horny part and immense rictus, reaching below the eyes, furnished with bristles. Rather large. Plumage soft and lax, much variegated … Caprimulgidæ.
Feet scarcely or not semipalmate, of frequently abnormal ratio of phalanges (middle or outer toe, or both, with fewer joints than usual among birds). Hallux very small, elevated, frequently lateral or versatile. Middle toe not produced nor its claw pectinate. Bill much as in the last, but rictus unbristled. Small. Plumage compact, of few simple subdued colors … Cypselidæ.
Family of PSITTACI
To characters of Psittaci add: Cere feathered, concealing the nostrils. Feet granular, rugose. Wings pointed. Tail cuneate. Plumage coarse and dry. Head feathered. Colors green, with yellow and blue … Psittacidæ.20
Analysis of Families of RAPTORES
Feet highly raptorial, with large, strong, sharp, curved, contractile claws, adapted for grasping. Hallux perfectly incumbent, lengthened (more than half as long as the fourth toe), with large claw. Front toes with slight basal webbing between outer or middle ones, or none; outer toe often reversible. Nostrils imperforate. Bill short, stout, not notably contracted in its continuity, with strongly hooked tip; tomia often once-twice toothed or lobed. Head feathered wholly or in greatest part. Lower larynx developed with one pair of muscles. Plumage with or without after-shafts. Cœca present, as a rule, if not always.
Physiognomy peculiar by reason of great lateral expansion and lengthwise shortening of the cranium, causing the eyes to be directed forward. Eyes surrounded by a disc of radiating bristly feathers, in front closely appressed to and hiding the base of the bill, elsewhere bounded by a rim of differently formed feathers. Tomia never toothed or lobed. Nostrils usually at the edge of the cere. Outer toe completely versatile, shorter than the inner toe. Basal phalanx of middle toe not longer than the second, and much shorter than the next. Legs commonly feathered or bristly to or on the toes. Plumage peculiarly soft and lax, without after-shafts; flight perfectly noiseless. Cranial walls widely separated by intervention of spongy diploë. Sternum commonly doubly notched. Chiefly nocturnal … Strigidæ.
Physiognomy not peculiar in any lateral expansion of the cranium; the eyes lateral in direction. No complete facial disc; base of bill not hidden by appressed bristles. Nostrils wholly in the cere. Outer toe rarely versatile, except Pandion, etc.; not shorter than the inner. Basal phalanx of middle toe longer than the second. Legs commonly naked and scutellate or reticulate in some portion of their length; toes always bare and scaly. Plumage compact, usually with after-shafts; flight audible. Cranial walls with little diploë. Sternum commonly single-notched or fenestrate, sometimes entire. Diurnal … Falconidæ.
Feet scarcely raptorial, with lengthened, little curved or contractile, weak, short claws. Hallux elevated, shortened, not more than half as long as the fourth toe, with small claw. Front toes all webbed at base; middle toe lengthened; outer not reversible. Basal phalanx of middle toe longer than either of the succeeding. Nostrils perforate. Bill lengthened and comparatively weak, little hooked, contracted in its continuity; tomia not toothed or lobed. Head naked of feathers in greatest part; sparsely bristly. No lower larnyx developed. No cœca. After-shafts absent … Cathartidæ.
Family of COLUMBÆ
With characters essentially as in Columbæ (exclusive of those peculiar to Diduncudidæ and Dididæ). Plumage without after-shafts; the feathers with thickened, spongy rhachis loosely inserted in the skin. Head small, completely feathered, excepting sometimes a circumorbital space. Tarsi naked or only feathered a little way above. Tail of twelve feathers, or lengthened, cuneate, and of fourteen. (Hallux not perfectly incumbent in Starnænas.) … Columbidæ.
Analysis of Families of GALLINÆ
Hind toe lengthened, insistent. Tail-feathers twelve. Sides of head and throat with naked spaces. Color greenish … Cracidæ.
Hind toe shortened, elevated. Tail-feathers usually fourteen or more. No green.
Large. Tarsi, toes, and nasal fossæ naked. Head bare of feathers, sparsely bristly, with wattles and caruncles. A pectoral tuft of bristly feathers. Tarsi usually spurred in the male. Plumage iridescent … Meleagrididæ.
Medium. Tarsi wholly or in great part, sometimes also the toes, and always the nasal fossæ, feathered. Head completely feathered, excepting a definite papillate strip over the eye. Tail-feathers sixteen or more. Sides of neck usually with lengthened feathers, or a naked distensible area, or both. No spurs. Plumage without iridescence … Tetraonidæ.
Small. Tarsi, toes, and nasal fossæ naked. Head completely feathered. No peculiar feathers or tympanum on sides of neck. No spurs. Plumage not iridescent … Perdicidæ.
Analysis of Families of LIMICOLÆ
Toes not lobate. Tarsi not notably compressed.
Legs extremely long; the tarsus equalling or exceeding the tail, and feet either four-toed and palmate (Recurvirostra), or three-toed and semipalmate (Himantopus); with the bill much longer than the head, very slender, acute, and curved upward … Recurvirostridæ.
Legs moderate, stout. Tarsus shorter than tail. Bill hard, more or less contracted at base, with short nasal fossa, gonydeal angle, and ascending gonys, the tip either compressed and truncate or depressed and acute. Feet three-toed and with basal webbing (Hæmatopus), or four-toed and cleft (Strepsilas) … Hæmatopodidæ.
Legs moderate. Tarsus shorter than tail, reticulate. Hind toe wanting (except in Squatarola, where very small, and in Aphriza). Bill short, straight,—not exceeding the head (generally shorter),—shaped like a pigeon’s, with short, broad, soft nasal fossæ separated by a constriction from the enlarged, obtuse, horny terminal part. Head large, globose, contracting suddenly to the bill. Neck short … Charadriidæ.
Legs moderate. Tarsus shorter than tail, scutellate. Hind toe present. Bill long,—equalling, or oftener exceeding, frequently several times longer than, the head; softish and membranous to the very tip, without constriction in its continuity; straight or variously curved … Scolopacidæ.
Toes lobate. Tarsi notably compressed.
General characters of Scolopacidæ. Body depressed; the under plumage thickened, duck-like. Habits natatorial … Phalaropodidæ.
Analysis of Families of HERODIONES
Hallux lengthened, perfectly incumbent, with large claw. Tarsi scutellate. Middle claw pectinate. Bill perfectly straight, tapering, acute. Loral region definitely naked, continuous with covering of the bill. Head narrow, elongate, tapering … Ardeidæ.
Hallux somewhat reduced, less perfectly incumbent. Tarsi commonly reticulate. Middle claw not pectinate. Lores, gular space and usually more of the head, naked. Bill variously curved or with expanded tip. (Genera Tantalus, Ibis, Mycteria, and Platalea.) … Tantalidæ.
Analysis of Families of ALECTORIDES
Of great stature, with extremely long neck and legs. Part or all of the head bare. Toes much shorter than the tarsi; with basal webbing, but without lobation; hallux very short, highly elevated. Bill equalling or exceeding the head, compressed, perfectly straight, contracted about the middle, with enlarged acute terminal portion; nasal fossæ wide and deep, with large perforate nostrils … Gruidæ.
Size moderate and small; neck and legs comparatively short. Head completely feathered, excepting, in the Coots and Gallinules, a broad horny frontal plate. Toes equalling or exceeding the tarsi, simple or lobate. Bill not constricted in the middle, rather shorter than the head, straight and quite stout; or much longer, regularly slender and decurved, with long nasal fossæ. Nostrils incompletely or not perforate … Rallidæ.
Analysis of Families of LAMELLIROSTRES
Of great stature, with extraordinarily lengthened neck and legs. Bill of unique shape, bent abruptly down from the middle. Tibiæ naked below. Hind toe minute or absent. Wings rounded. Red the chief color … Phœnicopteridæ.
Of moderate size; the neck short, or, when lengthened, not accompanied by co-ordinately lengthened legs, these being always shorter than the wing. Bill straight. Tibiæ feathered below. Hind toe present; well developed and functional, though short … Anatidæ.
[Pg xxviii] Analysis of Families of STEGANOPODES
Bill rather longer than head, cleft to eyes, very stout at base, tapering to the decurved, but not hooked, tip. Nostrils abortive. Gular sac moderate, naked. Wings rather long, pointed. Tail long, stiff, cuneate, twelve to fourteen feathered. Feet nearly beneath centre of equilibrium. General configuration goose-like … Sulidæ.
Bill several times longer than head, slender but strong, depressed, perfectly straight, with small distinct hooked nail at end. Nostrils very small. Gular sac enormous. Mandibular rami meeting only at tip. Wings extremely long, with upward of forty remiges. Tail short, rounded, of twenty or more feathers. Legs beneath centre of equilibrium, extremely short and stout … Pelicanidæ.
Bill about as long as head, stout, straight, scarcely tapering, strongly hooked. Nostrils abortive. Gular sac moderate, but evident; mostly naked. Wings short. Tail large, fan-shaped, scansorial, of twelve to fourteen broad stiff feathers, exposed to the base. Legs inserted far behind centre of equilibrium … Graculidæ.
Bill rather longer than head, slender, perfectly straight, tapering to an acute tip. Gular sac small. Nostrils minute. Wings and tail, and general configuration, as in the last … Plotidæ.
Bill much longer than head, straight, stout, strongly hooked. Nostrils very small. Gular sac well developed. Wings exceedingly long, strong, and pointed. Tail exceedingly long, deeply forked. Feet extraordinarily short; tarsi partly feathered … Tachypetidæ.
Bill about as long as head, straight, stout, tapering to an acute tip. Nostrils small. Gular sac rudimentary, feathered. Wings moderate, pointed. Tail short, but with two central feathers extraordinarily prolonged and filamentous. Feet small, beneath centre of equilibrium … Phæthontidæ.
Analysis of Families of LONGIPENNES
Nostrils not tubular, lateral, perforate. Bill with continuous covering, or only broken by a sort of cere, hooked or straight to the end. Hallux small and elevated, but always present … Laridæ.
Nostrils tubular, disjoined and lateral, or oftenest superior and united in one double-barrelled tube. Covering of bill in several pieces; bill always hooked. Hallux minute, rudimentary, or absent … Procellariidæ.
Analysis of Families of PYGOPODES
Feet four-toed, palmate. Hallux lobate, connected at base with base of inner toe. Tail perfect. Head closely and completely feathered. Nostrils with a depending lobe or flap. Bill straight, compressed, acute … Colymbidæ.
Feet four-toed, lobate. Hallux lobate, free. Tail rudimentary. Head with a naked loral strip and bristly or variously lengthened feathers. Nostrils simple. Bill straight or decurved at end, compressed, acute … Podicipidæ.
Feet three-toed, palmate. Hallux absent. Tail perfect. Head closely feathered or variously crested. Nostrils simple. Bill of indeterminate shape … Alcidæ.